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Full text of "Flora Indica Vol-I"

FLOHA INDICA.                                   239
carpels is a well known one, to which we only call attention as indicating an affinity
with Berberidea through Podophyllwm, with ZanZizabatea through Eolllollia nnd
all the typical genera of that Order, and with Papaveracea through Papaver itself,
which has broad placentso, and especially through the Mexican genus "RJomneya,., the
ovules of which are distributed over the whole cavity of the ovary. In- Cabonibece
the ovules are few, and confined to the dorsal suture of the carpels; and these are
free, indicating an affinity to Nelumbiacea on the one hand and Platyxtemon on the
other, a genus of Papaoeraceae with two free carpels
The seeds of NympJueacets are sometimes arillate, when the arillus forms an elon-
gated fleshy cup, arising from towards the base of the funiculus ^nd completely enve-
loping the seed. In most species the seeds arc completely imbedded in a cellular pulp
derived from the walls of the carpels and placental surfaces, affording a strong ana-
logy to the pulp si Lardisabalea and Podophyllum. The fact of the embryo being
enclosed in the nmniotic sac is well known to be common to this Order, and to some
very far removed from, it, as Piperacea and Sauntrea; but we have indicated a very
analogous structure in Moniiniacea, and we would further call attention to the strong
resemblance between the canal in the axis of the farinaceous albumen of Nymphaa-
cea and the cellular mass occupying the axis of the fleshy albumen of ffortonia,
and Boldoa. The relation of these to the amniotic sac is not made out, but we may
remark that they are certainly part of the nucleary sac of the ovule, and that in Hor-
tonia bat little albumen is developed in that part, which, remains cellular in the
ripe seed, whilst- in NymplxBa, owing to the cellular tissue itself being absorbed, an
open canal remains. The fact of the embryo lying in a cavity at the apes of the albu-
men, and not immersed in it, is repeated in Leontice and Bongardia, genera of Ber-
faridetp, where we have further indicated the sheath of the radicle as an important
modification of embryo-coverings, and requiring explanation.
Other peculiarities of flyMpfi/eacea, indicating their affinity, are that Cribombea
diflcr little from the ternary-sepaled RamtncuH> except fo the insertion, etc., of their
ovules and their amuiotic sac, and that they closely imitate in habit the Ranunculi
of the B&trachium section, The great disc of Nympluea is represented by that of
Ptvonia, as indicated by De CandoUe. In form the stigmata strongly resemble those
of'Papaver, as do the seeds to a great extent. The whorl of carpels of Nymphaa fur-
ther resembles in some degree that of IHieium and Dillenia, to which may be added
that Trecul describes the carpels of Nttphar as exhibiting a tendency to a dorsal dc-
hisccnce.                       *
We have thus a multitude of most important structural and physiological cha-
racters connecting Nymp7iaace<& with the Orders amongst which we place them, be-
sides many minor ones which are individually of little importance, but which together
establish an accumulation of affinities all pointing; in the same direction; to this we
may add, that we doubt if they agree with any other Natural Orders but the imme-
diate allies of these, in any characters of systematic importance.
Suborder I. NYMJPH WM.
Stamina plurima. Carpella in ovarium pluriloculare concreta. Qvula
plurima, parietibus ovarii undique affixa.
1. NYMPttflBA, L.
Sepala 4, imo toro inserta. Petala .12-20, 2-4-scriata. Stamina 40-
60, tnultiseriata. Ovariwn 6-84oculare; stigmatibus sessilibus linea-
ribus radiatis. Bacca spougiosa, irregulariter rupta. Semina in pnlpa
nidulantia, arillo sacciform! apico aperto induta; testa coriacea.
To any one who has studied a, numerous suite of specimens of tHe Indian species
of this beautiful genus, and the published descriptions of them, it will not be a matter
of surprise that we find it necessary to unite a considerable number*